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Costas completed his undergraduate studies at the Institute of Chemical and Bio-Engineering at the Swiss Federal Institute of Technology in Zurich (ETH Zurich, 2001) and his doctoral studies (Ph.D.) at the Department of Bioengineering at Imperial College London (IC, 2006) under the supervision of Danny O'Hare and Kim Parker. His Ph.D. thesis focussed on methods of nonstationary time-series analysis for electrochemical data. During his Ph.D. Costas was also a research scholar at the Department of Mathematics at the Massachusetts Institute of Technology (MIT) working with Martin Bazant on micro-/nano-scale phenomena occurring in the vicinity of charged surfaces. Since 2007, Costas is a postdoctoral fellow at the Division of Biology at the California Institute of Technology (Caltech) while maintaing affiliations with the Department of Bioengineering at IC and the Center of Molecular and Behavioural Neuroscience at Rutgers studying the effect of electric fields on the signalling characteristics (action potential initiation, synchronization, phase-locking, etc.) of single neurons and neuronal populations in collaboration with Christof Koch (Caltech), Mauricio Barahona (IC) and Gyuri Buzsaki (Rutgers).